Abstract
The niche-diversity hypothesis proposes that personality structure arises from the affordances of unique trait combinations within a society. It predicts that personality traits will be both more variable and differentiated in populations with more distinct social and ecological niches. Prior tests of this hypothesis in 55 nations suffered from potential confounds associated with differences in the measurement properties of personality scales across groups. Using psychometric methods for the approximation of cross-national measurement invariance, we tested the niche-diversity hypothesis in a sample of 115 nations (N = 685,089). We found that an index of niche diversity was robustly associated with lower intertrait covariance and greater personality dimensionality across nations but was not consistently related to trait variances. These findings generally bolster the core of the niche-diversity hypothesis, demonstrating the contingency of human personality structure on socioecological contexts.
Humans differ from one another in myriad ways—physically, behaviorally, and psychologically. Relatively stable patterns of individual differences in patterns of thinking, feeling, and behaving broadly define human personality (Briley & Tucker-Drob, 2014; Costa & McCrae, 1992a). Inductive methods, such as factor analysis, have characterized person-to-person variability in the broad constellation of personality-relevant traits into five or six major dimensions or factors (Digman, 1990; Lee & Ashton, 2004; Costa & McCrae, 1992b; Saucier & Goldberg, 1996). One dominant perspective holds that these factors, and how they vary and covary within populations, result directly from basic biological structures and processes that are intrinsic to the human mind (McCrae & Costa, 1997, 2008; Nettle, 2009). According to this universal-personality-structure view, the same patterns of personality-factor variation and covariation will be obtained across populations.
Contrasting with the predictions of the universal-personality-structure perspective, results from previous studies suggest that the number of dimensions needed to summarize personality variation differs somewhat across populations. Whereas differences in personality structure tend to be relatively small across Western and industrialized nations (Allik & McCrae, 2004; Kajonius & Mac Giolla, 2017; McCrae et al., 1998; Schmitt et al., 2007), inconsistencies are larger when low- and middle-income nations are considered (Heine & Buchtel, 2009; McCrae & Terracciano, 2005; Saucier et al., 2014). Moreover, research conducted with the Tsimane, a small-scale Amazonian society, suggests that two dimensions—prosociality and industriousness—are sufficient to characterize the major patterns of thinking, feeling, and behaving (Gurven et al., 2013). These Tsimane-specific dimensions each contain heterogeneous mixtures of items from the traditional five-factor traits (Gurven et al., 2013), making them qualitatively distinct from other two-factor models observed in industrialized societies (e.g., Digman, 1997).
Differences in personality structure between populations have been presumed by some researchers to stem from measurement artifacts, such as differences in translations or item functioning (Heine & Buchtel, 2009; McCrae & Terracciano, 2005). However, when such artifacts are controlled for, between-population differences in personality structure may also persist because of substantive differences in social and cultural dynamics underlying personality development (Briley & Tucker-Drob, 2017). The niche-diversity hypothesis (Lukaszewski et al., 2017; Smaldino et al., 2019) provides one framework for testing such an account.
The niche-diversity hypothesis holds that within human populations, individuals occupy different niches: micropopulations within a larger population with different affordances and cost–benefit structures (e.g., organizations, occupations, social and cultural groups, coalitions, families). For example, in human societies, some niches may favor high levels of patience and industriousness with low levels of anxiety but not specific levels of imaginativeness. Other niches may favor high levels of sociability and risk tolerance without specifically favoring any level of honesty. And some may not incentivize any particular levels or combinations of traits.
In general, humans tailor behavioral traits to the demands of local socioecological niches (Briley & Tucker-Drob, 2017; Henrich, 2015; Kaplan et al., 2009; Pinker, 2010; Tooby & DeVore, 1987). The developmental calibration of behavioral profiles may occur through a mixture of social learning (Legare, 2017) and state-behavior feedback loops (Sih et al., 2015). Consistent with the functionality of niche-behavior compatibility, research has shown that people whose multidimensional personality profiles are a better match with the demands of their niche experience higher material payoffs over time (Denissen et al., 2018).
The number of unique, specialized niches varies widely across human populations. In large complex societies, individuals can occupy an assortment of different niches—each incentivizing different optimum levels of personality traits. In smaller-scale societies, the number and diversity of niches that are available to be occupied—and the trait levels and combinations required for success within them—tend to be more delimited (Gurven et al., 2013). Thus, the number of niche-incentivized trait profiles is predicted to be greater in some human populations than others (Lukaszewski et al., 2017). Compared with populations with fewer unique niches, populations with more unique niches are hypothesized to exhibit (a) more distinct combinations of traits (i.e., less covariance among traits; Lukaszewski et al., 2017), (b) wider distributions of trait levels (i.e., more individual variation in each trait; Smaldino et al., 2019), and (c) more emergent dimensions of personality (Smaldino et al., 2019). Using agent-based models, Smaldino et al. confirmed that these predicted associations are obtained under conditions in which (a) populations differ in niche diversity and (b) individuals adapt trait levels on the basis of the niches that they occupy.
Statement of Relevance
Prominent models of personality structure posit that the patterning of thoughts, feelings, and behaviors is universal across human populations, resulting from innate dispositions that reliably create variation along five relatively distinct dimensions. But empirical work shows that there is much variation in personality structure across populations. What accounts for this variation? Recent theoretical developments suggest that personality structure can differ as a function of the number of micropopulations that incentivize different traits within a population (i.e., niches). Specifically, places with a greater number of diverse niches (e.g., workplaces, social groups) will exhibit less overlap in patterns of thoughts, feelings, and behaviors at the population level, producing more dimensions of personality than places with less niche diversity. We found support for this hypothesis using data from more than 680,000 people across 115 countries. These findings suggest that variation in the structure of personality is partially contingent on socioecological factors.
To date, only one data set of 55 nations (Schmitt et al., 2007) has been used to empirically test predictions from the niche-diversity hypothesis. In this sample, Lukaszewski et al. (2017) found a moderate negative association between nation-level intercorrelations among Big Five traits and a proxy of nation-level niche diversity based on three indices: the Human Development Index (HDI), the percentage of people living in cities (i.e., urbanization), and the variety of products produced within a nation (i.e., sectoral diversity). Using the same 55-nation sample and niche-diversity proxy, Smaldino et al. (2019) found that Big Five trait variance was moderately positively associated with niche diversity. Finally, Del Giudice (2021) applied dimensionality-reduction methods to the Big Five covariance estimates provided by Lukaszewski et al. (2017) for the 55 nations and found that niche diversity was positively associated with the number of emergent personality dimensions. These findings provide initial support for the niche-diversity hypothesis of personality structure.
Although prior studies implemented controls to assess artifacts of translations and general response tendencies (i.e., acquiescence bias), none have yet implemented methods to guard against the potential for the focal associations to arise from systematic differences in how the items within personality measures function across populations. The differential functioning of items across populations due to measurement nonequivalence can bias estimates of factor means, covariances, and variances (Byrne et al., 1989; Church et al., 2011). Thus, previous investigations of the association between niche diversity and personality structure derived from factor-score composites are potentially misleading.
Here, we present a stronger test of the niche-diversity hypothesis by drawing on new large-scale personality data from 115 nations and factor-analytic methods that approximate measurement invariance for a more valid comparison of personality structure across nations. We tested three focal predictions formalized by Smaldino et al. (2019): Niche diversity is (a) negatively associated with nation-level covariation among personality factors, (b) positively associated with nation-level variance across personality factors, and (c) positively associated with the number of nation-level personality dimensions.
Method
Participants
We drew personality data on 1,015,341 participants from the Open-Source Psychometrics Project (https://openpsychometrics.org/). The data sets from this repository have been used in a variety of publications (e.g., Chen et al., 2020; Hirschfeld et al., 2014; Kaiser et al., 2020). The data set contains responses to personality items along with Internet protocol (IP) addresses and country of residence; no information on participant sex or other demographics is available. Individuals accessed the survey through the Internet and received no compensation for participating. To minimize the potential for multiple instances from the same responders, we removed all participants with identical IP addresses. We also removed participants from nations with fewer than 90 participants. After these exclusions, our total sample size for analysis was 685,089 participants across 115 nations.
Measures
Personality
Personality was measured using the 50-item International Personality Item Pool five-factor markers (IPIP-FFM-50; Goldberg, 1992), which has strong construct validity (Lim & Ployhart, 2006) and is generally invariant across gender and American ethnic groups (Ehrhart et al., 2008). Sample items include, “I am the life of the party” (Extraversion), “I am interested in people” (Agreeableness), “I am always prepared” (Conscientiousness), “I am relaxed most of the time” (Emotional Stability), and “I have a rich vocabulary” (Openness). Participants rated the extent to which they agreed that each of the 50 statements was true of themselves using a 5-point scale (1 = disagree, 5 = agree). All instructions and scale items were presented in English to all participants. In our Results section, we present sensitivity analyses to test the role of English proficiency.
Niche diversity
We used the same index of niche diversity as did Lukaszewski et al. (2017) and Smaldino et al. (2019). A nation’s niche diversity is estimated as its score on the first principal component of three nation-level variables: sectoral diversity (i.e., the volume-weighted variety of a nation’s exports), urbanization (i.e., percentage of the nation’s population living in cities), and the HDI (i.e., a nation’s average levels of education, gross domestic product, and life expectancy). Each of these three variables is a conceptually appropriate indicator of a nation’s niche diversity, given their direct and indirect relationships to economic specialization and division of labor.
Sectoral diversity was supplied by Harvard University’s Atlas of Economic Complexity and is widely employed in macroeconomic modeling of country-level productivity and economic growth (https://atlas.cid.harvard.edu; Hausmann et al., 2014). A nation’s sectoral-diversity score is based on a volume-weighted estimate of its variety of exports—that is, how many different types of products a nation is able to produce (see Hausmann et al., 2014; Hidalgo & Hausmann, 2009). Diversity in exports by necessity requires specialization across firms producing goods for export. Sectoral-diversity scores therefore reflect division of labor, and thus economic-niche diversification, at the country level.
Urbanization and HDI variables were supplied by the United Nations. Urbanization is the percentage of the nation’s population living in urban (vs. rural) areas (United Nations Development Programme, 2020b). The HDI is a nation’s average levels of education, gross domestic product per capita, and life expectancy (United Nations Development Programme, 2020a). Urbanization and HDI reflect a nation’s niche diversity given (a) reciprocal relationships between the division of labor and the degree of urbanization at the national level (Gibbs & Martin, 1962) and (b) the interrelated effects of the division of labor and urbanization on a nation’s per-capita income and wealth. Division of labor and urbanization affect innovation and market productivity and efficiency (e.g., Henrich & Boyd, 2008; Rodriguez-Clare, 1996), which tends to fuel rising average incomes and standards of living (Hausmann et al., 2014; Hidalgo & Hausmann, 2009).
Of course, sectoral diversity, urbanization, and HDI are neither direct nor perfect measures of niche diversity. Each of these components of our niche-diversity composite potentially contains variance independent of niche diversity proper that can potentially drive any associations we find; however, they each also tap different conceptual aspects of niche diversity proper and contain nonoverlapping sources of confounding variance. Importantly, we conducted supplemental analyses to examine the focal relationships using each separate component of the niche-diversity index (see Section 3.1 in the supplementary materials at https://osf.io/7n4sr/), which showed the same pattern of results reported below.
For use in analyses, we pulled estimates of sectoral diversity, urbanization, and HDI for the year 2015, which is the year before the Open-Source Psychometrics Project began collecting personality data. Because data were missing on one or more of these indices for several nations, we conducted a principal component analysis that allows for missing data (Dray & Dufour, 2007). The first principal component of these three indicators explained 80% of the variance in the data, and loadings on this first principal component were strong for each indicator (HDI = .80, urbanization = .73, sectoral diversity = .82). Higher scores on this z-scaled index suggest a greater number of specialized niches, whereas lower scores suggest fewer specialized niches within a nation. Figure 1 shows each nation’s estimated niche diversity in relation to its sample size.

Map highlighting the nations and geographic regions represented by the personality data used in the current study. The centroid size represents the relative sample size for each nation; nations with smaller points contributed fewer participants. The hue and luminance of each centroid depict that nation’s standing on the niche-diversity index; nations with lower scores on the niche-diversity index are lighter, and those with higher scores are darker.
Analytic methods
We used a two-step analytic procedure. In the first step, we specified the five-factor structure with the IPIP-FFM-50 items modeled in the context of a multiple-group model, using the alignment method to maximize approximate measurement invariance across groups (i.e., nations). The alignment allowed for meaningful comparison of latent-factor means, variances, and covariances across groups without imposing untenable assumptions regarding strict equivalence of factor loadings and intercepts (Asparouhov & Muthén, 2014). In the second step, we used metaregression to examine the relationship of niche complexity to the covariance and variance parameter estimates extracted from the first step—and their associated standard errors—while controlling for several potential confounds. Details of each method are described in their respective sections below.
Multigroup alignment
Our research question required that we compare variances and covariances across nations. It is therefore necessary to address differences in how the personality items function across nations (Byrne et al., 1989; Church et al., 2011). We addressed measurement invariance using the alignment method (Asparouhov & Muthén, 2014). Assessments of measurement invariance become intractable with many groups (Marsh et al., 2018). Asparouhov and Muthén (2014) developed the multigroup-alignment method as an automated alternative that approximates measurement invariance, allowing for comparison of groups without requiring strict invariance across groups. The multigroup-alignment method uses a two-step procedure.
In the first step, the configural model is estimated on the basis of the researcher-specified factor structure in which all loadings and intercepts are freely estimated; factor variances are fixed to one and means to zero. Because the alignment method cannot handle cross-loadings or be combined with exploratory structural equation modeling (ESEM) approaches, we assumed the simple Big Five factor structure with no cross-loadings. Typically, omitted cross-loadings will inform the factor covariance estimate, so the factor covariance estimates that we obtained should not be interpreted literally as differences in the relations between real dimensions of trait variation but, instead, as a more general indication that the constellation of items underlying each set of personality factors differ in their overall magnitude of association with one another. Considering that complex factor structures with many items—such as the IPIP-FFM-50 used here—are unlikely to meet common fit thresholds (cf. Marsh et al., 2010), the simple Big Five configural model exhibited reasonable fit in most nations (root mean square error of approximation [RMSEA]: M = .07, minimum = .05, maximum = .09; for full results of the confirmatory factor analysis for each nation, see Section 3.3 in the supplementary materials at https://osf.io/7n4sr/). To examine the degree to which deviations from the simple factor structure may influence our results, we present sensitivity analysis in nations with acceptable RMSEA values (i.e., < .08) in the Results section.
In the second step of the alignment procedure, all factor variances and means are iteratively estimated across groups using a loss-simplicity function that minimizes the noninvariance and provides the most invariant pattern across groups. This loss-simplicity function is analogous to different factor rotations in exploratory factor analysis in that the aligned model will have the same fit as the configural model. We used the fixed alignment method, which identifies group-factor means and variances in relation to a fixed group whose respective mean and variance remain fixed at zero and one. We opted to set the United States as the reference group to facilitate comparison with the broader literature, most of which is based on American participants. 1
The approximation of measurement invariance provided by the multigroup alignment method is intended to provide more accurate comparison of covariances, variances, and means across groups (Asparouhov & Muthén, 2014). Our alignment results revealed generally poor invariance across nations (average invariance index = .33), which is itself consistent with the niche-diversity model’s prediction that personality-factor structure is not uniform across nations. Although this suggests that the alignment method is likely an imperfect solution to the problem of measurement invariance, it still represents an improvement over previous approaches using simple sum scores that do nothing to ameliorate measurement error and bias. Simulations demonstrate that the alignment method accurately recovers known population parameters even under conditions of substantial noninvariance across items, especially when sample sizes are large, as they generally are in the current sample (Asparouhov & Muthén, 2014).
Metaregression
The multigroup-alignment method does not currently offer support for estimating correlates of group-specific parameters, so examination of the focal associations between niche diversity and the covariance and variance estimates requires additional analytic steps. To examine the focal relationships of niche diversity to personality covariance and variance, we used the metaregression methods described by Tucker-Drob et al. (2019). We constructed separate models for each of the two focal meta-analytic outcomes: (a) nation-level interfactor covariance estimates and (b) nation-level intrafactor variance estimates. In the interfactor covariance models, the absolute values of each of the 10 parameter estimates for the pairwise covariances among Big Five factors across the 115 nations were modeled as the meta-analytic outcome. In the intrafactor variance models, the five parameter estimates for each of the Big Five factor variances across the 115 nations were modeled as the meta-analytic outcome. In all primary models, we clustered these parameter estimates by nation. We also added precision weights proportional to the inverse-sampling variance of the parameter estimates, so that samples with more precise estimates were weighted more strongly in the metaregression analysis.
Effective dimensionality
To examine the focal relationship between niche diversity and the number of emergent personality dimensions in each nation, we implemented the effective-dimensionality technique described by Del Giudice (2021). Effective dimensionality provides an estimate of the dimensionality of a set of variables, much like exploratory factor analysis; however, effective dimensionality provides a continuous, nondiscrete estimate of dimensionality and is agnostic about the underlying causal structure of the variables, making it superior to selecting a discrete number of factors on the basis of an arbitrary eigenvalue cutoff. There are several indices of effective dimensionality based on different derivations; we employed the n1 index because it does not assign disproportional weight to larger eigenvalues, making it suitable for general-purpose estimations of effective dimensionality (for a review, see Del Giudice, 2021).
We used the R code provided by Del Giudice (2021) to estimate the effective dimensionality of the Big Five factors in each nation. Importantly, rather than using items as input, we constructed covariance matrices using the interfactor covariances and the intrafactor variance estimates obtained from the alignment analyses as input for the effective-dimensionality analysis, which provides the added benefits of approximate measurement equivalence. We addressed small-sample bias using Mestre’s (2008) method (for details, see Del Giudice, 2021).
Analytic procedure
We conducted the alignment and metaregression analyses in Mplus (Version 7.1.4; Muthén & Muthén, 2012). We carried out all other aspects of data cleaning, analyses, and data visualization in the R programming environment (Version 4.0.2; R Core Team, 2020). All data and code used to conduct the analyses and create the figures presented in this article, along with the supplementary materials, are provided on OSF (https://osf.io/7n4sr/).
In our primary metaregression analyses, we regressed each meta-analytic outcome (i.e., Big Five interfactor covariances and intrafactor variances) on the index of niche diversity. In our primary effective-dimensionality analyses, we regressed each nation’s effective-dimensionality estimate on the index of niche diversity. In secondary analyses, we examined the robustness of the focal associations to alternative explanations and potential confounds by including 11 nation-level controls in our analyses: mortality and homicide rates, acquiescence bias, the Big Five factor means, English-proficiency estimates, sample size, and geographic region. We describe each control variable and the rationale for its inclusion below. Figure 2 presents the intercorrelations among these nation-level control variables.

Intercorrelations among nation-level niche-diversity estimates and nation-level covariates. The personality controls labeled for each Big Five factor (Agreeableness, Conscientiousness, Extraversion, Emotional Stability, and Openness) refer to the alignment-estimated latent-factor means for each nation. Nation-level acquiescence was calculated as the mean acquiescence score across participants within each nation. Sample size and homicide rate were log-transformed. The size of each oval indicates the magnitude of the association (thinner = stronger, thicker = weaker), and the direction of the oval’s tilt indicates the direction of the association (right tilt = positive, left tilt = negative). HDI = Human Development Index.
Mortality and homicide rates
Med-edović (2020) recently proposed that the previously reported differences in personality structure across nations (Lukaszewski et al., 2017; Smaldino et al., 2019) may be driven not by niche diversity but, rather, by differences in life-history-related behavioral diversification resulting from cross-population variation in exposure to environmental harshness. Although “environmental harshness” can be interpreted in multiple ways (Stearns & Rodrigues, 2020), we included two commonly used nation-level proxies to test against this alternative explanation: homicide rates and mortality rates. We obtained homicide rates from the United Nations Office on Drugs and Crime (n.d.). We log-transformed the homicide-rate variable for analyses because it was positively skewed. We computed each nation’s mortality rate as the mean of adult male and adult female mortality rates for the year 2015 obtained from The World Bank (2019a, 2019b). For nations in which data were not available for the year 2015, we imputed the estimate from the closest preceding or succeeding year. No mortality estimates were present for any year for Taiwan, so we imputed the mortality rate from neighboring China.
Acquiescence bias
Acquiescence bias refers to the tendency of participants to be more inclined to agree with items than they are to disagree (or vice versa), potentially confounding variance and covariance estimates. We calculated acquiescence bias according to methods reported by Soto et al. (2008) by computing the mean for each participant across pairs of items from each IPIP-FFM-50 factor subscale with opposite valence before reverse coding the items. The opposite-valence item pairs from each subscale are as follows: “I don’t like to draw attention to myself” and “I don’t mind being the center of attention” from the Extraversion subscale, “I often feel blue” and “I seldom feel blue” from the Emotional Stability subscale, “I am not really interested in others” and “I am interested in people” from the Agreeableness subscale, “I like order” and “I leave my belongings around” from the Conscientiousness subscale, and “I do not have a good imagination” and “I have a vivid imagination” from the Openness subscale. Higher numbers on the resulting acquiescence score represent a tendency to acquiesce (“yea-saying”), whereas lower numbers represent a tendency to dissent (“nay-saying”). We averaged the acquiescence score across participants within each nation to estimate nation-level acquiescence scores.
Big Five latent-factor means
We included the latent nation means for each of the Big Five factors estimated by the alignment-parameter estimates to control for any nation-level differences in personality trait levels that may influence responses to personality scales (e.g., evaluative bias; Lukaszewski et al., 2017), potentially biasing the nation-level trait covariance and variance estimates.
English proficiency
Because the personality survey was administered only in English, a selection bias exists where nations with lower English proficiency were selected against. Participants from nations with lower English proficiency have a higher potential to misunderstand the questions, creating bias. To assess whether English proficiency could be driving any observed effects, we conducted a sensitivity analysis using only the nations in which English is an official language (either de facto or de jure official).
We also included nation-level estimates of English proficiency provided by Education First (2021) as a control in our robustness analyses. For nations where there were no English-proficiency data from the year 2015, we imputed the closest available preceding or succeeding estimate. English-proficiency data were completely unavailable for 27 nations, so we imputed either (a) the highest available English proficiency estimate for the 21 nations where proficiency data were missing but English is an official language or (b) the mean English-proficiency estimate for six nations where English is not an official language and proficiency data were missing.
Sample size
Lukaszewski et al. (2017) included sample size as a control to demonstrate that the differences in accuracy of covariance estimates were not driving the relationship to niche diversity. Our metaregression analyses with inverse-variance weights already accounted for the influence of sample size on the covariance and variance estimates, and our measure of effective dimensionality also corrected for sample size bias. Still, we included sample size as a control variable in robustness analyses because the sample size contributed by each nation likely serves as a proxy for potentially unmeasured confounds related to differential selection. We log-transformed this sample-size variable because it was positively skewed.
Geographic region
We included dummy-coded variables representing the continent on which each nation is located to control for nonindependence of observations due to geospatial proximity.
Results
Primary analyses
Across nations, the mean interfactor covariance estimate was .18 (SD = .02), the mean intrafactor variance estimate was .86 (SD = .03), and the mean effective-dimensionality estimate was 4.47 (SD = .32). Interfactor covariance estimates were positively correlated with intrafactor variance estimates (r = .52) and negatively correlated with effective-dimensionality estimates (r = −.78). Effective-dimensionality estimates were negatively correlated with intrafactor variance estimates (r = −.35).
Figure 3 shows the results of the primary metaregression analyses examining overall interfactor covariance, intrafactor variance, and effective dimensionality as a function of niche diversity. 2 Each standard-deviation increase in niche diversity was associated with .52 standard deviations lower covariance among the Big Five factors; in terms of raw units, each standard-deviation increase in niche diversity was associated with .02 lower interfactor covariance. The association between niche diversity and variance within the Big Five factors was not statistically different from zero. Finally, each standard-deviation increase in niche diversity was associated with a .53-standard-deviation increase in effective dimensionality, or in raw units, each standard-deviation increase in niche diversity was associated with an average of .13 more effective dimensions of personality. This pattern of results was obtained using each of the individual predictors of niche diversity as well (see Section 3.1 in the supplementary materials).

Bubble plots depicting absolute overall interfactor covariance (top), overall intrafactor variance (middle), and effective dimensionality (bottom) of the Big Five traits across 115 nations as a function of the nations’ standing on the niche-diversity index. Nations are grouped according to five geographic regions. The size of each bubble in the intrafactor variance and interfactor covariance plots is proportional to the inverse-sampling variance weight given to the estimates for each country; larger circles indicate greater weight in the analysis as a result of smaller standard errors and less sampling variance. Lines show best-fitting regressions (solid lines indicate significant regressions; the dashed line indicates a nonsignificant regression), and error bands show 95% confidence intervals. CI = confidence interval.
Sensitivity checks
Because the use of absolute values of the covariance estimates in our analyses may have biased estimates, we also conducted analyses in which we did not force all estimates to take on positive values. We reversed the covariance estimates for pairs that were on average negative and left any negative estimates untouched for pairs where the mean covariance was positive. Thus, for any given pair of variables, covariances could take on both positive and negative values. Metaregression results were not appreciably different from the analyses using absolute values of the covariance estimates (β = −0.48, p = .001, 95% confidence interval [CI] = [−0.74, −0.21]). To disentangle any potential confounding effects of personality variance on personality covariance, we conducted analyses of interfactor correlations, rather than covariances, by scaling the interfactor covariances and their standard errors relative to their respective intrafactor variances. The association with niche diversity was slightly stronger than when using covariances (β = −0.69, p < .001, 95% CI = [−0.92, −0.46]).
There was more variability in the residual interfactor covariance and effective-dimensionality estimates at low levels of niche diversity than at high levels. We therefore examined the focal relationships in only the 84 countries with niche-diversity scores greater than zero, for which the residuals were more homoscedastic. The negative interfactor covariance association (β = −0.41, p < .001, 95% CI = [−0.61, −0.21]) and the positive effective-dimensionality associations (β = 0.43, p < .001, 95% CI = [0.24, 0.61]) were both robust to these sensitivity checks. Restricting our analyses to the 42 nations in the sample with English as an official language did not substantively change the associations between niche diversity and interfactor covariance (β = −0.41, p = .003, 95% CI = [−0.68, −0.14]), intrafactor variance (β = 0.01, p = .961, 95% CI = [−0.41, 0.43]), or effective dimensionality (β = 0.52, p < .001, 95% CI = [0.24, 0.79]). Finally, in analyses including only the 90 nations that exhibited reasonable fit in confirmatory factor analyses (i.e., RMSEA < .08), the associations were qualitatively unchanged between niche diversity and interfactor covariances (β = −0.51, p < .001, 95% CI = [−0.69, −0.32]), intrafactor variances (β = 0.18, p = .100, 95% CI = [−0.03, 0.39]), and effective dimensionality (β = 0.43, p < .001, 95% CI = [0.29, 0.56]).
Robustness analyses
We conducted secondary metaregression analyses to examine the robustness of the focal relationships to potential confounds. We added the sample size for each nation, nation-level estimates of acquiescence bias, alignment-estimated latent-factor means for each of the Big Five traits for each nation (i.e., evaluative bias), English proficiency, homicide rate, mortality rate, and dummy codes representing the macrogeographic regions as controls for geospatial dependence to the primary models. Figure 4 shows the results of the robustness tests, and we provide tabular output with exact p values for all parameters in the supplementary materials (Section 3.1).

Point estimates for focal associations with overall absolute interfactor covariance (left column), overall intrafactor variance (middle column), and effective dimensionality (right column) after including controls, along with point estimates for the nation-level control variables. The reference group for the continent dummy variable is Europe. The personality controls labeled for each Big Five factor refer to the alignment-estimated latent-factor means for each nation. Sample size and homicide rate were log-transformed. Error bars represent 95% confidence intervals.
As shown in Figure 4, when all controls were included as predictors, niche diversity was still negatively associated with Big Five interfactor covariances (β = −0.52, p < .001, 95% CI = [−0.75, −0.28]) and positively associated with effective dimensionality (β = 0.41, p < .001, 95% CI = [0.20, 0.62]). The association between niche diversity and intrafactor variance became negative with the inclusion of controls but remained nonsignificant (β = −0.25, p = .095, 95% CI = [−0.55, 0.04]). Nation-level acquiescence bias and Conscientiousness were significantly negatively associated with interfactor covariances and significantly positively associated with effective dimensionality across nations (for effect sizes and CIs, see Fig. 4). Nation-level Extraversion was also significantly negatively associated with effective dimensionality and significantly positively associated with interfactor covariance (for effect sizes and CIs, see Fig. 4). Intrafactor variance across nations was significantly positively associated with nations’ sample size and average Openness and significantly negatively related to nations’ acquiescence bias (for effect sizes and CIs, see Fig. 4). We also examined the effects of the different sets of control variables by conducting separate analyses, entering the controls in a stepwise manner; these results are provided in the supplementary materials (Section 3.1).
Exploratory analyses of pairwise covariance and individual variances
Figure 5 shows the associations (with and without controls) between niche diversity and each of the 10 pairwise absolute covariance estimates (left panel) and each individual intrafactor variance estimate (right panel). One of the 10 pairs of Big Five interfactor covariances was significantly associated with niche diversity in the same direction in both the controlled and uncontrolled analyses after correction for false-discovery rate: Conscientiousness with Extraversion (ps < .008). No other pairwise factor covariances were significantly associated with niche diversity in the same direction both with and without controls. None of the associations between niche diversity and individual Big Five intrafactor variances were statistically significant and in the same direction in analyses with and without controls.

Point estimates for the associations between niche diversity and the absolute interfactor covariances among all possible pairs of Big Five traits (left) and each of the individual Big Five intrafactor variances (right). Results are shown separately for models including and not including the 11 control variables. Error bars represent 95% confidence intervals. A = Agreeableness; C = Conscientiousness; E = Extraversion; ES = Emotional Stability; O = Openness.
Discussion
The niche-diversity hypothesis proposes that personality structure is a function of the number of socioecological niches within a population (Lukaszewski et al., 2017; Smaldino et al., 2019). It predicts that increasing niche diversity is associated with (a) lower personality trait covariance, (b) greater personality trait variance, and (c) greater personality dimensionality. We improved on the only prior empirical test of this hypothesis by more than doubling the number of nations under investigation and implementing methods to alleviate confounds associated with differences in the psychometric properties of personality scales across nations. In a sample of 685,089 individuals across 115 nations, we found robust evidence for the first and third predictions but little overall support for the second.
As predicted, we found that personality traits were less distinct in nations with lower niche diversity. Not only was this association obtained using a psychometric method to minimize artifacts associated with measurement inequivalence across nations, but the association remained after models statistically controlled for several potential confounds. Importantly, niche diversity explained variation in personality structure in the presence of environmental-harshness indicators, which casts doubt on an alternative explanation proposed by Med-edović (2020) that population differences in personality structure may be accounted for by the effects of environmental harshness on behavioral diversification along the life-history spectrum.
We did not find robust support for the model-predicted relationship between niche diversity and overall trait variance. This association was not statistically significant and reversed when models controlled for potential confounds. This finding indicates that the association between trait variance and niche diversity within human populations may not be as straightforward as prior simulations and data suggested (Smaldino et al., 2019) and reveals where further refinement of the model may be fruitful. Specifically, niche diversity may not inevitably lead to greater trait variance if many niches within and between populations tend to incentivize similar levels of traits when they are incentivized at all. If this were the case, the diversity of niche-incentivized trait combinations could still vary across populations, leading to higher personality dimensionality and less overall covariance among traits, as we found.
One limitation of this study is that the personality survey was administered only in English via the Internet. This enabled us to use identical personality measures across all participants and nations, avoiding confounds associated with translation. It does mean, however, that many of the participants responded to a survey in their nonnative language and that individuals who were not English literate were unable to participate. To guard against this potential language confound, our metaregression models adjusted for variation in acquiescent responding across nations and variation in national estimates of English proficiency. Importantly, selection bias associated with both English literacy and Internet access may render our results more conservative if participants tended to come from more urban, niche-diverse regions of their countries. Nonetheless, personality assessments administered in participants’ preferred languages, allowing more representative sampling, will be necessary to fully assess the generalizability of these effects.
Another limitation is that our data lack demographic information about participants. Prior research suggests that the factor structure of personality may vary with age (Beck et al., 2019; Mõttus et al., 2019; Soto et al., 2008). If the age distribution of participants within each nation systematically varied with niche diversity, then the results presented here could be confounded with age trends in the personality structure. Although we do not have a strong reason to suspect such confounding, the lack of demographic data prevents us from exploring this issue here. However, the niche-diversity hypothesis may partly explain age trends in personality structure. For instance, observed age differences in personality structure among U.S. participants (Beck et al., 2019) appear to track trends of workforce participation and thereby occupational-niche diversification. Examining the extent to which age-related differences in personality structure coincide with age differences in niche diversity is an interesting future direction.
Finally, our analysis of personality structure relied on the Big Five factor structure even though the niche-diversity hypothesis explicitly assumes that the structure of personality varies across cultures. Because it was not possible to combine the alignment with ESEM approaches, we chose alignment to prioritize maximizing the comparability of personality traits across countries. To quote Cronbach and Meehl (1955), we used the simple five-factor model as a tool for “defining a working reference frame, located in a convenient manner” rather than to discern “‘real dimensions’ [in which] a great deal of surplus meaning is implied” (pp. 288 and 287, respectively). Research investigating the degree to which complex factor solutions and other personality frameworks yield different patterns of association with niche diversity may yield additional insights into cultural variation in personality structure.
The niche-diversity hypothesis was partly motivated by recognizing that personality science could benefit from model building from first principles (Lukaszewski et al., 2020). Our findings provide the strongest support to date for the central empirical predictions generated by the niche-diversity hypothesis, demonstrating the contingency of personality structure on socioecological dynamics. Further considerations of the socioecological factors that vary within and between human societies will be crucial for refining our understanding of the nature of personality and psychological variation more generally.
Footnotes
Acknowledgements
We thank Courtney Crosby, Anna Sedlacek, Steven Gangestad, William Revelle, and an anonymous reviewer for their constructive feedback on this article.
Transparency
Action Editor: Steven W. Gangestad
Editor: Patricia J. Bauer
Author Contributions
P. K. Durkee identified the data source. P. K. Durkee developed the analysis plan with E. M. Tucker-Drob and conducted the analyses under the supervision of E. M. Tucker-Drob. P. K. Durkee drafted the manuscript, and E. M. Tucker-Drob, A. W. Lukaszewski, C. R. von Rueden, M. D. Gurven, and D. M. Buss contributed critical revisions. All the authors approved the final manuscript for submission.
