Regional palaeoenvironmental reconstructions and data on artiodactyl response to climate change suggest that large game densities would have expanded in response to increasingly mesic conditions during the late Holocene in the Wyoming Basin. We use the prey model of foraging theory to predict late-Holocene increases in artiodactyls, relative to lagomorphs and rodents, and more specifically in bison relative to pronghorn. This prediction is then tested against 284 dated archaeofaunas from the Wyoming Basin. Close fits are found between the deductively derived prediction and the empirical records. Although artiodactyls demonstrate a general increasing trend across the most recent 5000 years, during the period of most intense human occupation hunting pressure appears to have depressed large game populations.
Ahlbrandt, T.S.1974: Dune stratigraphy, archaeology, and chronology of the Killpecker dune field. In Wilson, M. , editor, Applied geology and archaeology: the Holocene history of Wyoming . Geologic Survey of Wyoming, 51—60.
2.
Bailey, J.A.1990: Management of Rocky Mountain Bighorn sheep herds in Colorado . Special Report Number 66. Colorado Division of Wildlife .
3.
Bayham, F., and Hatch, P.1985: Archaeofaunal remains from the New River area. In Doyel, D.E. and Elson M.D., editors, Hohokam settlement and economic systems in the Central New River Drainage, Arizona. Soil Systems Publications in Archaeology Number 4, 405—33.
4.
Beiswenger, J.M.1991: Late Quaternary vegetational history of Grays Lake, Idaho . Ecological Monographs61, 156—82.
Broughton, J.M.1994: Declines in mammalian foraging efficiency during the Late Holocene, San Francisco Bay. Journal of Anthropological Archaeology13, 371—401.
7.
——— 1999: Resource depression and intensification during the late Holocene, San Francisco Bay: evidence from the Emeryville Shellmound vertebrate fauna. Anthropological Records32. University of California Publications.
8.
——— 2002: Prey spatial structure and behavior affect archaeological tests of optimal foraging models: examples from the Emeryville Shellmound vertebrate fauna. World Archaeology34, 60—83.
9.
Broughton, J.M. and Bayham, F.E.2003: Showing off, foraging models, and the ascendance of large game hunting in the California Middle Archaic. American Antiquity6, 783—89.
10.
Brown, A.L. and Litvatis, J.A.1995: Habitat features associated with predation of New England cottontails: what scale is appropriate?Canadian Journal of Zoology73, 450—59.
11.
Brown, C.F. and Krausman, P.R.2003: Habitat characteristics of 3 leporid species in southeastern Arizona. Journal ofWildlife Management67, 83—89.
12.
Brown, D.E., Fagan, W.F., Lee, R., Shaw, H.G. and Turner, B.2003: Winter precipitation and Pronghorn fawn survival . Arizona Antelope Foundation. Retrieved 26 September 2007 from http://www.azantelope.org/Facts___Research/Winter_Precipitation___Fawn_Su/winter_precipitation___fawn_su.html
13.
Bryson, R.A.1989: Late Quaternary volcanic modulation of Milankovitch climate forcing. Theoretical and AppliedClimatology39, 115—25.
14.
——— 1992: A macrophysical model of the Holocene intertropical convergence and jetstream positions and rainfall for the Saharan region. Meteorology and Atmospheric Physics47, 247—58.
15.
Bryson, R.A. and Bryson, R.U.1997: Macrophysical climate modeling of Africa's late Quaternary climate: site-specific, high-resolution applications for archaeology . African Archaeological Review14, 143—60.
16.
Bryson, R.U. and Bryson, R.A.1998: Application of a global volcanicity time-series on high-resolution paleoclimatic modeling of the eastern Mediterranean. In Issar, A.S. and Brown, N., editors, Water, environment and society in times of climate change. Kluwer Academic Publishers, 1—19.
17.
——— 2000: The modeled climate history of Green River, Wyoming: since the last glacial maximum. In Pastor, J.V. , Thompson, K.W., Talbot, R.K., Eckerle, W.P. and Ingbar, E.E., editors, `Seeds-Kee-Dee' riverine adaptation in southwest Wyoming. Museum of Peoples and Cultures Technical Series No. 99—3, Brigham Young University, Appendix F.
18.
Buechner, H.K.1950: Range ecology of pronghorn on the Wichita Mountains Wildlife Refuge. Transactions of the North American Wildlife Conference15, 627—44.
19.
Byers, D.A. and Broughton, J.M.2004: Holocene environmental change, artiodactyl abundances, and human hunting strategies in the Great Basin. American Antiquity69, 251—71.
20.
Byers, D.A. and Ugan, A.2005: Should we expect large game specialization in the late Pleistocene? An optimal foraging perspective on early Paleoindian prey choice . Journal of Archaeological Science32, 1624—40.
21.
Byers, D.A., Smith C. and Broughton, J.M.2005: Holocene artiodactyl population histories and large game hunting in the Wyoming Basin, USA. Journal of Archaeological Science32, 125—42.
22.
Byers, J.A. and Hogg, J.T.1995: Environmental effects on prenatal growth rate in pronghorn and bighorn: further evidence for energy constraint on sex-biased maternal expenditure. BehavioralEcology4, 51—57.
23.
Cannon, M.D.2000: Large mammal relative abundance in pithouse and Pueblo Period archaeofaunas from southwestern New Mexico: resource depression among the Mimbres-Mogollon?Journal of Anthropological Archaeology19, 317—47.
24.
——— 2003: A model of central place forager prey choice and an application to faunal remains from the Mimbres Valley, New Mexico. Journal of Anthropological Archaeology22, 1—25.
25.
Carrera, P.E., Short, S.K. and Shroba, R.R.1985: A pollen study of Holocene peat and lake sediments, Leidy Peak area, Uinta Mountains, Utah. GeologyStudies32, 1—7.
26.
Chamrad, A.D. and Box, T.W.1965: Drought-associated mortality of range grasses in south Texas. Ecology46, 780—85.
27.
Chapman, J.A. and Willner, G.R.1978: Sylvilagus audubonii. Mammalian Species106, 1—4.
28.
DeBloois, D.L., Cranney, S.J., Blackwell, B., Bates, B., Flinders, S., Grandison, J. and Cornicelli, L.2001: Utah big game annual report 2001. Publication Number 01—30 . Utah Division of Natural Resources Division of Wildlife Resources.
29.
Douglas, C.L.2001: Weather, disease and bighorn lamb survival during 23 years in Canyonlands National Park. Wildlife SocietyBulletin29, 297—305.
30.
Eckerle, W.P.1997: Eolian geoarchaeology of the Wyoming Basin. In Larsen, M.L. and Francis, J., editors, Changing perspectives on the archaic of the Northwest Plains. University of South Dakota Press , 138—67.
31.
Eckerle, W.P., Hobey, J., and Francis, J.E.1999: The environmental and cultural setting. In Miller, M.E., Sunders, P.H. and Francis, J.E., editors, The Trappers Point Site (48SU1006): early Archaic adaptations in the Upper Green River Basin. Cultural Resource Series No. 1. Office of the Wyoming State Archaeologist, University of Wyoming , 5—22.
32.
Eckerle, W.P., Hanna, R. and Taddie, S.2002: Geoarchaeological assessment of the Blue Point Site 48SW5734, Sweetwater County, Wyoming: Alberta-Cody Complex adaptation to the Green River Basin uplands. In Johnson, D. and Pastor, J., editors, The Blue Point Site: Paleoindian/Archaic transition in Southwest Wyoming. Western Archaeological Services, Appendix A.
33.
Evans, G., Westphal, V., Welch, M., Karpowski, J., Grandison, J., Cornicelli, L., Giunta, B. and Cranney, S.J.1997: Utah big game annual report 1997. Publication Number 97—17 .Utah Division of Natural Resources Division of Wildlife Resources.
34.
Fairbanks, S.W.1993: Birthdate, birthweight, and survival in pronghorn fawns . Journal of Mammology74,129—35.
35.
Fenneman, N.M.1931: Physiography of Western United States. McGraw-Hill Book Company, Inc.
36.
Forman, S.L., Oglesby, R. and Webb, R.S.2001: Temporal and spatial patterns of Holocene dune activity on the Great Plains of North America: megadroughts and climate links . Global and Planetary Change29, 1—29.
37.
Fox, L.M., Krausman, P.R., Morrison, M.L. and Katting, R.M.2000: Water and nutrient content of forage in Sonoran pronghorn habitat, Arizona. California Fish and Game88, 216—32.
38.
Frank, D.A. and McNaughton, S.J.1992: The ecology of plants, large mammalian herbivores and drought in Yellowstone National Park. Ecology73, 2043—58.
39.
Frolik, A.L. and Keim, F.D.1933: Native vegetation in the Prairie Hay District of north central Nebraska. Ecology14, 298—305.
40.
Grayson, D.K.1977: Paleoclimatic implications of the Dirty Shame Rockshelter mammalian fauna. Tebiwa9, 1—26.
——— 1987: The biogeographic history of small mammals in the Great Basin: observations on the past 20,000 years. Journal of Mammology68, 359—75.
43.
——— 1991: Alpine faunas from the White Mountains, California: adaptive change in the Late Prehistoric Great Basin?Journal of Archaeological Science18, 483—506.
44.
——— 2000a: The Homestead Cave mammals. In Madsen, D.B., editor, Late Quaternary paleoecology in the Bonneville Basin. Bulletin130. Utah Geological Survey, 67—90.
45.
——— 2000b: Mammalian responses to middle Holocene climatic change in the Great Basin of the western United States. Journal of Biogeography27, 181—92.
46.
Hawkes, K.1991: Showing off: tests of a hypothesis about men's hunting goals. Ethology and Sociobiology12, 29—54.
47.
——— 1993: Why hunter-gatherers work: an ancient version of the problem of public goods. Current Anthropology34, 341—61.
48.
Henrickson, L.S.2004: Frozen bison and fur trapper's journals: building a prey choice model for Idaho's Snake River Plain. Journal of Archaeological Science31, 903—16.
49.
Hildebrandt, W.R. and Jones, T.L.2002: Depletion of prehistoric pinniped populations along the California and Oregon Coasts: were humans the cause? In Kay, C.E. and Simmons, R.T., editors, Wilderness and political ecology: Aboriginal influences and the original state of nature. University of Utah Press, 72—110.
50.
Hildebrant, W.R. and McGuire, K.R.2002: The ascendance of hunting during the California Middle Archaic: an evolutionary perspective. American Antiquity68, 790—92.
51.
Howell, J.A., Brooks, G.C., Semenoff-Irving, M. and Greene, C.2002: Population dynamics of tule elk at Point Reyes National Seashore, California. Journal ofWildlife Management66, 478—90.
52.
Hunter, M.D. and Price, P.W.1992: Playing chutes and ladders: heterogeneity and the relative roles of bottom-up and top-down forces in natural communities. Ecology73, 724—32.
53.
Janetski, J.C.1997: Fremont hunting and resource intensification in the eastern Great Basin. Journal of Archaeological Science24, 1075—88.
54.
Jense, G.K., Burruss, J.W., Bates, J.W., Bowden, N.I., Coles, F., Kimball, J.F., Pederson, J.C. and Nielson, R.J.1979: Utah big game investigations and management recommendations 1978—1979. Publication Number 79—6. Utah Department of Fish and Game.
55.
Jense, G.K., Fitzgerald, J.W., Bates, J.W. and Bowden, N.I.1984: Utah big game investigations and management recommendations 1983—1984. Publication Number 84—2. Utah Department of Fish and Game.
56.
John, R.T., Bowden, N.I., Bates, J.W., Boss, A., Coles, F., Kimball, J.C. and Nielson, R.J.1974: Utah big game investigations and management recommendations 1973—1974. Publication number 74-5. Utah Department of Fish and Game.
57.
Kitchen, D.W. and O'Gara, B.W.1982: Pronghorn. In Chapman, J.A. and Feldhamer, G.A., editors, Wild animals of North America. Johns Hopkins University Press, 960—71.
58.
Knight, D.H.1994: Mountains and plains: the ecology of Wyoming landscapes . Yale University Press.
59.
Larsen, M.L.1997: Housepits and mobile hunter-gatherers: a consideration of the Wyoming evidence. Plains Anthropologist42, 353—69.
60.
Larsen, M.L. and Francis, J.1997: Changing perspectives on the Archaic of the Northwest Plains. University of South Dakota Press.
61.
Legler, R.P.1970: Habitat preference of the desert cottontail with additional notes on the black-tailed jack rabbit. Unpublished masters thesis . New Mexico State University.
62.
Leslie, D.M. and Douglas, C.L.1979: Desert bighorn sheep of the River Mountains, Nevada . Wildlife Monographs66,1—57.
63.
Li, X.R., Jia, X.H. and Dong, G.R.2006: Influence of desertification on vegetation pattern variations on the cold semi-arid grasslands of Qinghai-Tibet Plateau, north-west China . Journal of Arid Environments64, 505—22.
Longhurst, W.M., Connolly, G.E., Browning, B.M. and Garton, E.O.1979: Food interrelationships of deer and sheep in parts of Mendocino and Lake Counties, California. Hilgardia47, 191—247.
66.
Lubinski, P.M.1997: Pronghorn intensification in the Wyoming Basin: a study of mortality patterns and prehistoric hunting strategies. Unpublished Ph.D. dissertation. Department of Anthropology, University of Wisconsin.
67.
——— 2000: Of bison and lesser mammals: prehistoric hunting patterns in the Wyoming Basin. In Madsen, D.B. and Metcalf, M.D., editors, Intermountain archaeology. University of Utah Anthropological Papers Number 122. University of Utah Press, 179—88.
68.
Lupo, K.D. and Schmitt, D.N.1997: On late Holocene variability in bison populations in the northeastern Great Basin. Journal of California and Great BasinAnthropology19, 50—69.
69.
Martner, B.E.1986: Wyoming climate atlas. University of Nebraska. Mayer, J.2002: Evaluating natural site formation processes in eolian dune sands: a case study from the Krmpotich Folsom Site, Killpecker dunes, Wyoming, Journal of Archaeological Science29, 1199—211.
70.
——— 2004: Late quaternary stratigraphy and geochronology of the western Killpecker dunes, Wyoming, USA. Quaternary Research61, 72—84.
71.
McCartney, C.W. and Miller, M.W.1998: Modeling the population dynamics of bighorn sheep: a synthesis of the literature. Special Report Number 73. Colorado Division of Wildlife.
72.
McCollough, Y.B.1980: Niche separation of seven North American ungulates on the National Bison Range, Montana. Ph.D. dissertation, University of Michigan.
——— 1978: Bison. In Schmidt, J.L. and Gilbert, D.L., editors, Big game of North America: ecology and management. Stackpole Books, 123—33.
75.
Miller, J.C.1992: Geology in archaeology: geology, paleoclimates, and archaeology in the Western Wyoming Basin. Unpublished Masters thesis, University of Wyoming.
76.
Monroe, J.S.2003: Holocene timberline and palaeoclimate of the Northern Uinta Mountains, Northeastern Utah, USA. The Holocene13, 175—86.
77.
Moss, J.H.1951: Early man in the Eden Valley. Museum Monographs No. 6. University Museum, University of Pennsylvania.
78.
Murphy, A.H.1970: Predicted forage yield based on fall precipitation in California annual grasslands. Journal of Range Management23, 363—65.
Peek, J.M., Dennis, B. and Hershey, T.2002: Predicting population trends of mule deer. Journal ofWildlife Management66, 729—36.
81.
Picton, H.D.1984: Climate and the prediction of reproduction of three ungulate species. Journal of Applied Ecology21, 869—79.
82.
Power, M.E.1992: Top-down and bottom-up forces in food webs: do plants have primacy?Ecology73, 733—46.
83.
Quirt-Booth, T. and Cruz-Uribe, K.1997: Analysis of leporid remains from prehistoric Sinagua Sites, northern Arizona. Journal of Archaeological Science24, 945—60.
84.
Ripple, W.J. and Beschta, R.L.2004: Wolves, elk, willows and the trophic cascades in the upper Gallatin Range of southeastern Montana, USA. ForestEcology and Management200, 161—81.
85.
Schlesinger, W.H., Reynolds, J.F., Cunningham, G.L., Huenneke, L.F., Jarrell, W.M., Virginia, R.A. and Whitford, W.G.1989: Biological feedbacks in global desertification . Science247, 1043—48.
86.
Schmitt, D.N., Madsen, D.B. and Lupo, K.D.2002: Small-mammal data on early and middle Holocene climates and biotic communities in the Bonneville Basin, USA. Quaternary Research58, 255—60.
87.
Singer, F.C., Harting, A., Symonds, K.K. and Coughenour, M.B.1997: Density dependence, compensation, and environmental effects on elk calf mortality in Yellowstone National Park. Journal ofWildlife Management61, 12—25.
88.
Smith, C.S.1988: Seeds, weeds, and prehistoric hunters and gatherers: the plant macrofossil evidence from southwest Wyoming. Plains Anthropologist33, 141—58.
89.
——— 2003: Hunter-gatherer mobility, storage, and houses in a marginal environment: an example from the mid-Holocene of Wyoming. Journal of Anthropological Archaeology22, 162—89.
90.
Smith, C.S. and McNees, L.M.1999: Facilities and hunter-gatherer long-term land use patterns: an example from southwest Wyoming. American Antiquity64, 117—36.
91.
Smith, C.S., Martin, W. and Johansen, K.A.2001: Sego lilies and prehistoric foragers: return rates, pit ovens, and carbohydrates. Journal of Archaeological Science28, 169—83.
92.
Smith, N.S. and Krausman, P.R.1988: Desert bighorn sheep: a guide to selected management practices . Biological Report85(35). United States Fish and Wildlife Service .
93.
Stapley, H.D., Bates, J.W., Henderson, R.S., Huff, C.L., Nielson, R., Nish, D. and Winn, J.1966: Utah big game investigations and management recommendations 1965—1966. Publication Number 66—6. Utah Department of Fish and Game.
94.
Stephens, D.W. and Krebs, J.R.1986: Foraging theory. Princeton University Press.
95.
Stephenson, T.E., Holecheck, J.L. and Kuykendall, C.B.1985: Drought effect on pronghorn and other ungulate diets. Journal ofWildlife Management49, 146—51.
96.
Stokes, S. and Gaylord, D.R.1993: Optical dating of holocene dune sands in the Ferris Dune Field, Wyoming. Quaternary Research39, 274—81.
97.
Stuiver, M. and Reimer, P.J.2004: Calib 14C calibration program home page. Retrieved 26 September 2007 from http://calib.qub.ac.ukSzuter, C.R. and Bayham, F.E.1989: Sedentism and prehistoric animal procurement among desert horticulturalists of the North American Southwest . In Kent, S., editor, Farmers as hunters: the implications of sedentism. Cambridge University Press, 80—95.
98.
Ugan, A.2005: Does size matter? Body size, mass collecting, and their implications for understanding prehistoric foraging behavior. American Antiquity70, 75—90.
99.
Van Dyke, W.A., Sands, A., Yoakum, J., Polenz, A. and Blaisdell, J.1983: Wildlife habitats inmManaged rangelands — the Great Basin of southeastern Oregon: bighorn sheep. General Technical Report PNW-159 . Pacific Northwest Forest and Range Experiment Station, United States Forest Service.
100.
Van Vuren, D. and Bray, M.P.1986: Population dynamics of bison in the Henry Mountains, Utah . Journal of Mammalogy67, 503—11.
101.
White, T.R.C.1978: The importance of relative shortage of food in animal ecology. Oecologia33, 71—86.
102.
Whitlock, C.1993: Postglacial vegetation and climate of Grand Teton and southern Yellowstone National Parks. Ecological Monographs63, 173—98.
103.
Whitlock, C. and Bartlein, P.1993: Spatial variations of Holocene climate change in the Yellowstone region. Quaternary Research39, 231—38.
104.
Whitlock, C., Bartlein, P. and Van Norman, K.J.1995: Stability of climate regimes in the Yellowstone region. Quaternary Research43, 433—36.
105.
Wiessner, P.2002: Hunting, healing, and hxaro exchange: a long-term perspective on !Kung (Ju/'hoansi) large-game hunting. Evolution andHuman Behavior23, 407—36.
106.
Wolverton, S.2005: The effects of the Hypsithermal on prehistoric foraging in Missouri. American Antiquity70, 91—106.
107.
Wydeven, A.P. and Dahlgren, R.B.1985: Ungulate habitat relationships in Wind Cave National Park. Journal ofWildlife Management49, 805—13.
108.
Yoakum, J.D.2004: Foraging ecology, diet studies and nutrient values . In O'Gara, B.W. and Yoakum, J.D., editors, Pronghorn: ecology and management. University Press of Colorado, 447—502.
109.
Zhang, J., Zhao, H., Zhang, T., Zhao, X. and Drake, S.2005: Community succession along a chronosequence of vegetation restoration on sand dunes in Horqin Sandy Land. Journal of Arid Environments62, 555—66.
